Basic Perks Edit
This perk is extremely useful if combat speed is a bottleneck. This perk will not affect the time it takes to move a cell after killing an enemy, nor will it affect the time between one group of trimp dying and another starting the fight (if there is one; needs verification).
The perk is considerably resources is a bottleneck, or if the time it takes to breed up a new army is significant. This perk will not affect you negatively even in that case.
In the long grind between level 50 and 59, where your trimps are not dying and battle takes up a long time, each upgrade roughly increases your clear speed by 5% multiplicatively. And 20 upgrades will multiply your clear speed by roughly 2.8.
Convenient at the beginning of the game, and slightly useful during the short period in the early mid-game when a significant fraction of your trimps are dying very fast, and your nurseries/venimps haven't completely kicked in yet. Not very useful, and becomes less so with pheromones. But a few levels means you don't have to trap trimps 1 by 1 early game.
Useful early game; becomes insignificant really quickly. Not a particularly good perk. Does stack with Carpentry however. Early-game, it is a good idea to get lots of levels in this and then respec out of it later on - you will keep the bonus on all the Territory Bonuses you've gotten so far.
Trimp breeding is linaer. This upgrade increases the base speed of breeding. Extremely potent if trimp breeding is a limiting factor. Superb upgrade to get if you suffer a significant down time due to slowness of breeding. Not so useful if you don't.
This is a convenience upgrade. The benefit of this is to decrease the relative cost of storage upgrades compared to the amount stored. With no upgrade, it takes 1/4 of the resource to double storage. As such, if you want enough storage for 8 hours of production, you have to wait 1 hour. With 5 levels in this, it takes 1/8 of the resource to double storage. So you only have to wait half an hour if you want storage for 8 hours.
Increases your production, useful perk.
Increases your clear speed; kind of like agility. In fact, it does almost the exact thing as agility if trimp breeding is not an issue. Linear increase, however, compared to compounding increase of agility. At higher levels, agility is more useful.
Increases your health. Useful upgrade from 1 to 50. Makes it easier to survive one shot from quick attacking enemies, and gives more time for your trimps to breed. Not so useful from 50-59, as you take no damage, and your block value is significantly higher than your hp.
Useful as a direct multiplier to helium gained (except those from dimension of anger). Not that useful for other things (except maybe a bit of gem and fragments early on). The gain from production quickly makes the gain from looting insignificant.